Your bridge isn't bridging properly. If Router B is sending a destination unreachable then the packets are being handled on it further up the stack at layer 3 by some sort of routing component rather than by a layer 2 bridging one.
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Indeed it is not. Do you, by chance, have any suggestions -- troubleshooting, alternatives, etc.?
run ip route
and ip route get $CLIENT_PUBLIC_IP
on router B and see if it has a route to the client, and/or if the default route is correct. Its default gateway might not be set correctly (it should be router A)
and responds appropriately (SYN, ACK),
Does it respond to the client address (public IP?)
I'm not exactly sure what the previous issue was, but it appears that, possibly, the previous bridge that was in use was broken in some way. I have since switched the primary router to one that supports WDS, and created a WDS bridge between the two, and now everything is working as expected.